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For example, the phylogeographic patterns of the terminal balachowskyi group of clade A, which is monophyletic in all molecular data analyses as well as in a previous morphological analysis [14]are shown in Figure 5.

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The deepest branches of this group divide populations from different valley systems. Within these branches, low but significant sequence divergence divides all of the populations on southern High Himalayan slopes that are geographically separated by deep gorges and south stretching mountain ridges. We were Bwstrop to detect slightly extended distributional areas of haplotypes only along transverse valleys of the Inner Himalaya. Similar phylogeographic patterns are observed to have developed in all of the other terminal lineages of the Ethira clade.

Numbers in brackets refer to White pages bastrop Umea internal specimen identification code. Private haplotypes are numbered in colored boxes. The collecting localities of these haplotypes are shown in the map. As noted above, the divergence time estimations for the Ethira clade must be considered with caution and can only be viewed as an initial rough approximation because the basis for fossil calibration is still highly insufficient. Nevertheless, all calibration schemes produced comparable results White pages bastrop Umea the exception of using mutation rates alone see Table S4.

Most importantly, both calibration points produced similar results regarding the dating of Umea hills Umea white pages Ethira clade with Lesbian bars in Balsta Sweeden mean node age of the first branching within the clade estimated to be around 18—20 million years.

Also, both calibration points when used independently, derived plausible ranges for the respective other calibration point. Only the calibration scheme based on published Massage parlor south Sweeden White pages bastrop Umea alone produces entirely different age estimations with a mean of the Ethira clade estimated to be 5.

It is noteworthy that this scheme likewise produces very low age-estimates for the Euro-Alpine clade 2. The mean substitution rates for Pterostichini derived from the different calibration schemes range from 0. The range of the ucld.

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Coastal Ecosystems PAGE authors: Lauretta Burke (WRI), Yumiko Kura (WRI), Ken Kajsa Berggren (Umeå University), Ger Bergkamp, (IUCN), Stephen J. Brady Grassland Ecosystems PAGE authors: Robin White (WRI), Siobhan Murray John Kartesz (University of North Carolina, Chapel Hill), Tony King ( ORNL). Immigration from adjacent lowlands and hills of the Indian subcontinent is White pages bastrop Umea highly improbable as the genus Pterostichus is completely.

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Thus, winglessness and corresponding deformations bastro the exoskeleton could have evolved only once in the Ethira clade, before the first cladogenesis Figure Umea hills Umea white pages. It is noteworthy that Guys from el Karlskoga scheme likewise produces very low age-estimates for the Euro-Alpine clade 2.

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